La nucleación: Una alternativa para la restauración ecológica de bosques neotropicales

Autores/as

  • Julián E. Díaz-Triana Departamento de Biología, Facultad de Ciencias, Universidad Nacional de Colombia. Bogotá, Colombia
  • Orlando Vargas-Ríos Departamento de Biología, Facultad de Ciencias, Universidad Nacional de Colombia. Bogotá, Colombia
  • Nelly Rodríguez-Eraso Departamento de Biología, Facultad de Ciencias, Universidad Nacional de Colombia. Bogotá, Colombia

DOI:

https://doi.org/10.25260/EA.23.33.3.0.2134

Palabras clave:

experiencias de nucleación, Neotrópico, nucleación aplicada, nucleación complementaria, gestión adaptativa en paisajes, sucesión por parches, técnicas de nucleación

Resumen

La deforestación y la degradación de tierras debido al uso intensivo del suelo son amenazas predominantes para los bosques neotropicales. En este marco, la restauración ecológica (RE) se plantea como una alternativa para contrarrestar la pérdida de biodiversidad y de servicios ecosistémicos, lo cual continúa siendo un reto y una prioridad global. La nucleación es una técnica que busca formar microhábitats que faciliten la regeneración natural y amplíen las interacciones entre organismos, promoviendo la sucesión ecológica. Aunque la formación de islas de árboles (nucleación aplicada) se usa a menudo por su relación costo-efectividad en comparación con otras alternativas de RE, su implementación en bosques neotropicales enfrenta desafíos que van desde la definición de objetivos precisos hasta su escalamiento a grandes áreas. En nuestro trabajo presentamos un panorama de las alternativas de nucleación para restaurar áreas de bosques tropicales, y evaluamos 1) sus fundamentos, condiciones y beneficios; 2) los estudios teóricos y experimentales, y 3) los desafíos para aplicarla en diferentes escenarios de restauración. Observamos que la nucleación es un modelo de restauración adaptable a escenarios con variabilidad en el potencial de recuperación y que abarca múltiples objetivos mediante diversas técnicas. Pese a que los estudios se han centrado en comparar resultados a corto plazo entre la nucleación aplicada y otras estrategias, la composición y la selección de especies, las configuraciones alternativas para la combinación de técnicas y el escalamiento a grandes áreas son temas que permanecen poco abordados. Sugerimos la implementación y el monitoreo de la nucleación deben orientarse hacia la experimentación dentro de los proyectos, y que la investigación futura debería considerar tratamientos alternativos con miras a una gestión adaptativa.

Citas

Abilash, P. C. 2021. Restoring the Unrestored: Strategies for Restoring Global Land during the UN Decade on Ecosystem Restoration (UN-DER). Land 10:2-19. https://doi.org/10.3390/land10020201.

Aguilar-Garavito, M. 2014. Técnicas y estrategias de restauración ecológica. Pp. 142-171 en M. Cabrera and W. Ramírez (eds.). Restauración ecológica de los páramos de Colombia. Transformación y herramientas para su conservación. Instituto de Investigación de Recursos Biológicos Alexander von Humboldt (IAvH), Bogotá, D.C., Colombia.

Albornoz, F. E., A. Gaxiola, B. J. Seaman, F. I. Pugnaire, and J. J. Armesto. 2013. Nucleation-driven regeneration promotes post-fire recovery in a Chilean temperate forest. Plant Ecology 214:765-776. https://doi.org/10.1007/s11258-013-0206-x.

Armenteras, D., and N. Rodríguez. 2014. Dinámicas y causas de deforestación en bosques de Latinoamérica: una revisión desde 1990. Colombia Forestal 17(2): 233-246. https://doi.org/10.14483/udistrital.jour.colomb.for.2014.2.a07.

Badano, E. I., O. R. Samour-Nieva, J. Flores, J. L. Flores-Flores, J. A. Flores-Cano, and J. P. Rodas-Ortíz. 2016. Facilitation by nurse plants contributes to vegetation recovery in human-disturbed desert ecosystems. Plant Ecology 9(5):485-497. https://doi.org/10.1093/jpe/rtw002.

Balvanera, P. 2012. Los servicios ecosistémicos que ofrecen los bosques tropicales. Ecosistemas 21(1-2):136-147. URL: revistaecosistemas.net/articulo.asp?Id=709.

Barrera‐Cataño, J. I., J. Garibello, C. Moreno‐Cárdenas, and S. Basto. 2022. Trade-offs at applying tree nucleation to restore degraded high andean forests in Colombia. Restoration Ecology 31(3):e13753. https://doi.org/10.1111/rec.13753.

Bechara, F. C., S. J. Dickens, E. C. Farrer, L. Larios, E. N. Spotswood, P. Mariotte, and K. N. Suding. 2016. Neotropical rainforest restoration: comparing passive, plantation and nucleation approaches. Biodiversity and Conservation 25:2021–2034. https://doi.org/10.1007/s10531-016-1186-7.

Bechara, F. C., B. E. Trentin, V. Lex Engel, D. A. Estevan, and T. Ticktin. 2021. Performance and cost of applied nucleation versus high-diversity plantations for tropical forest restoration. Forest Ecology and Management 491:119088. https://doi.org/10.1016/j.foreco.2021.119088.

Bhadouria, R., R. Sing, P. Srivastava, and A. S. Raghubanshi. 2016. Understanding the ecology of tree-seedling growth in dry tropical environment: a management perspective. Energy, Ecology and Environment 1(5):296-309. https://doi.org/10.1007/s40974-016-0038-3.

Boanares, D., and C. S. de Azevedo. 2014. The use of nucleation techniques to restore the environment: A bibliometric analysis. Natureza and Conservação 12: 93-98. https://doi.org/10.1016/j.ncon.2014.09.002.

Carlucci, M. B., L. da S. Duarte, and V. D. Pillar. 2011. Nurse rocks influence forest expansion over native grassland in southern Brazil. Journal of Vegetation Science 22:111-119. https://doi.org/10.1111/j.1654-1103.2010.01229.x.

Casalla, R., and J. Korb. 2018. Termite diversity in Neotropical dry forest of Colombia and the potential role of rainfall in structuring termite diversity. Biotropica 51:165-177. https://doi.org/10.1111/btp.12626.

Cayuela, L., and I. Granzow-de la Cerda. 2012. Biodiversidad y conservación de bosques neotropicales. Ecosistemas 21(1-2):1-5. URL: revistaecosistemas.net/articulo.asp?Id=725.

Ceccon, E. 2013. Restauración de bosques tropicales: fundamentos ecológicos, prácticos y sociales. Editorial Díaz De Santos. México, D.F., México.

Cole, R. J., K. D. Holl, and R. A. Zahawi. 2010. Seed rain under tree islands planted to restore degraded lands in a tropical agricultural landscape. Ecological Applications 20:1255-1269. https://doi.org/10.1890/09-0714.1.

Connel, J., and R. Slatyer. 1977. Mechanims of succession in natural communities and their role in community stability and organization. American Naturalist 111:1119-1144. https://doi.org/10.1086/283241.

Corbin, J. D., and K. D. Holl. 2012. Applied nucleation as a forest restoration strategy. Forest Ecology and Management 265:37-46. https://doi.org/10.1016/j.foreco.2011.10.013.

Corbin, J. D., G. R. Robinson, L. M. Hafkemeyer, and S. N. Handel. 2016. A long-term evaluation of applied nucleation as a strategy to facilitate forest restoration. Ecological Applications 26:104–114. https://doi.org/10.1890/15-0075.

Csákvári, E., Z. Molnár, and M. Halassy. 2022. Estimates of regeneration potential in the Pannonian sand region help prioritize ecological restoration interventions. Communications Biology 5:1136. https://doi.org/10.1038/s42003-022-04047-8.

Derroire, G., R. Coe, and J. R. Healey. 2016. Isolated trees as nuclei of regeneration in tropical pastures: testing the importance of niche-based and landscape factors. Journal of Vegetation Science 27:679-691. https://doi.org/10.1111/jvs.12404.

Díaz-Gallegos, J. R., and J. F. Mas-Caussel. 2009. La deforestación de los bosques tropicales: una revisión. Mapping 136:83-96.

Díaz-Triana, J. E., S. Torres-Rodríguez, L. Muñoz-P, and A. Avella-M. 2019. Monitoreo de la restauración ecológica en un bosque seco tropical interandino (Huila, Colombia): programa y resultados preliminares. Caldasia 41(1):43-30. https://doi.org/10.15446/caldasia.v41n1.71318.

Dimson, M., and T. W. Gillespie. 2020. Trends in active restoration of tropical dry forest: Methods, metrics, and outcomes. Forest Ecology and Management 467:118150. https://doi.org/10.1016/j.foreco.2020.118150.

FAO and PNUMA. 2020. El estado de los bosques del mundo 2020. Los bosques, la biodiversidad y las personas. El Estado del Mundo. ONU, Roma, Italia. https://doi.org/10.4060/ca8642es.

Filho, N. L., G. Rodrigues dos Santos, and R. L. Ferreira. 2013. Comparando técnicas de nucleação utilizadas na restauração de áreas degradadas na amazonia brasileira. Revista Árvore, Viçosa-MG 37(4):587-597. https://doi.org/10.1590/S0100-67622013000400002.

Franks, S. J. 2003. Facilitation in multiple life-history stages: Evidence for nucleated succession in coastal dunes. Plant Ecology 168:1-11. https://doi.org/10.1023/A:1024426608966.

Fremout, T., E. Thomas, H. Taedoumg, S. Briers, C. E. Gutiérrez-Miranda, et al. 2021. Diversity for Restoration (D4R): Guiding the selection of tree species and seed sources for climate-resilient restoration of tropical forest landscapes. Journal of Applied Ecology 59:664-679. https://doi.org/10.1111/1365-2664.14079.

Gann, G. D., T. McDonald, B. Walder, J. Aronson, C. R. Nelson, J. Jonson, J. G. Hallett, C. Eisenberg, M. R. Guariguata, J. Liu, F. Hua, C. Echeverría, E. Gonzales, N. Shaw, K. Decleer, and K. W. Dixon. 2019. International principles and standards for the practice of ecological restoration. Second edition. Restoration Ecology 27:S1-S46. https://doi.org/10.1111/rec.13035.

García-Barrios, L., Y. M. Galván-Miyoshi, I. A. Valdivieso-Pérez, O. R. Masera, G. Bocco, and J. Vandermeer. 2009. Neotropical Forest Conservation, Agricultural Intensification, and Rural Out-migration: The Mexican Experience. BioScience 59(10):863-873. https://doi.org/10.1525/bio.2009.59.10.8.

Garibello, J., L. Riaño, J. Cuellar, J. I. Barrera-Cataño, and W. Ramírez. 2021. Identificación de vacíos de investigación aplicada para restaurar ecosistemas terrestres en Colombia. Colombia Forestal 24(1):88-107. https://doi.org/10.14483/2256201X.15679.

Gómez-Aparicio, L., R. Zamora, J. M. Gómez, J. A. Hódar, J. Castro, and E. Baraza. 2004. Applying plant facilitation to forest restoration: a meta-analysis of the use of shrubs as nurse plants. Ecological Applications 14(4):1128-1138. https://doi.org/10.1890/03-5084.

Griscom, H. P., and M. S. Ashton. 2011. Restoration of dry tropical forest in Central America: A review of pattern and process. Forest Ecology and Management 261:1564-1579. https://doi.org/10.1016/j.foreco.2010.08.027.

Grubb, P. J. 1977. The maintenance of species-richness in plant communities: the importance of the regeneration niche. Biological Reviews 52(1):107-145. https://doi.org/10.1111/j.1469-185X.1977.tb01347.x.

Hartshorn, G. S. 2002. Biogeografía de los bosques neotropicales. Pp. 59-82 en M. R. Guariguata and G. H. Kattan (eds.). Ecología y conservación de bosques neotropicales. Editorial Libro Universitario Regional, Cartago, Costa Rica. ISBN 9968-801-11-9.

Herrera-Varón, Y., and M. Pinzón-Arias. 2021. Generalidades del monitoreo participativo y su aplicación en la restauración ecológica. Pp. 64-73 en M. Aguilar-Garavito and W. Ramírez (eds.). Evaluación y seguimiento de la restauración ecológica en el páramo andino. Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Bogotá D. C., Colombia.

Herrick, J. E., J. W. van Zee, S. E. McCord, E. M. Courtright, J. W. Karl, et al. 2015. Monitoring Manual for Grassland, Shrubland, and Savanna Ecosystems Volume 1: Core Methods. Second Edition. USDA-ARS Jornada Experimental Range, Las Cruces, New México, USA.

Holl, K. D. 2017. Restoring tropical forest form the bottom up. Science 355:455-456. https://doi.org/10.1126/science.aam5432.

Holl, K. D., J. C. Lesage, T. Adams, J. Rusk, R. D. Schreiber, and M. Tang. 2021. Vegetative spread is key to applied nucleation success in non-native-dominated grasslands. Restoration Ecology 29:1-9. https://doi.org/10.1111/rec.13330.

Holl, K. D., J. L. Reid, J. M. Chaves-Fallas, F. Oviedo-Brenes, and R. A. Zahawi. 2017. Local tropical forest restoration strategies affect tree recruitment more strongly than does landscape forest cover. Journal of Applied Ecology 54:1091-1099. https://doi.org/10.1111/1365-2664.12814.

Holl, K. D., J. L. Reid, R. J. Cole, F. Oviedo-Brenes, J. A. Rosales, and R. A. Zahawi. 2020. Applied nucleation facilitates tropical forest recovery: Lessons learned from a 15-year study. Journal of Applied Ecology 57:2316–2328. https://doi.org/10.1111/1365-2664.13684.

Holl, K. D., V. M. Stout, J. L. Reid, and R. A. Zahawi. 2013. Testing heterogeneity-diversity relationships in tropical forest restoration. Oecologia 173:569-578. https://doi.org/10.1007/s00442-013-2632-9.

Holl, K. D., R. A. Zahawi, R. J. Cole, R. Ostertag, and S. Cordell. 2011. Planting Seedlings in Tree Islands Versus Plantations as a Large-Scale Tropical Forest Restoration Strategy. Restoration Ecology 19:470-479. https://doi.org/10.1111/j.1526-100X.2010.00674.x.

Howe, H. F., and C. Martínez-Garza. 2014. Restoration as experiment. Botanical Sciences 92(4):459-468. https://doi.org/10.17129/botsci.146.

Hulvey, K. B., E. A. Leger, L. M. Porensky, L. M. Roche, K. E. Veblen, A. Fund, J. Shaw, and E. S. Gornish. 2017. Restoration islands: a tool for efficiently restoring dryland ecosystems? Restoration Ecology 25:S124–S134. https://doi.org/10.1111/rec.12614.

Hunter, O. M., M. Keller, D. Morton, B. Cook, M. Lefsky, M. Ducey, S. Saleska, R. C. de Oliveira, and J. Schietti. 2015. Structural dynamics of tropical moist forest gaps. PLoS ONE 10(7):e0132144. https://doi.org/10.1371/journal.pone.0132144.

Kazemi, H., H. Klug, and B. Kamkar. 2018. New services and roles of biodiversity in modern agroecosystems: A review. Ecological Indicators 93:1126-1135. https://doi.org/10.1016/j.ecolind.2018.06.018.

Lambin, E. F., P. Meyfroidt, X. Rueda, A. Blackman, J. Börner, et al. 2014. Effectiveness and synergies of policy instruments for land use 527 governance in tropical regions. Global Environmental Change 28:129-140. https://doi.org/10.1016/j.gloenvcha.2014.06.007.

Lindig-Cisneros, R. 2017. Ecología de Restauración y Restauración Ambiental. UNAM. México, D. F., México. https://doi.org/10.22201/enesmorelia.9786070294525e.2017.

Löf, M., A. Bolte, D. F. Jacobs, and A. M. Jensen. 2014. Nurse Trees as a Forest Restoration Tool for Mixed Plantations: Effects on Competing Vegetation and Performance in Target Tree Species. Restoration Ecology 22(6):758-765. https://doi.org/10.1111/rec.12136.

Martínez-Garza, C., F. Bongers, and L. Poorter. 2013. Are functional traits good predictors of species performance in restoration plantings in tropical abandoned pastures? Forest Ecology and Management 303:35-45. https://doi.org/10.1016/j.foreco.2013.03.046.

Méndez-Toribio, M., C. Martínez-Garza, and E. Ceccon. 2021. Challenges during the execution, results, and monitoring phases of ecological restoration: Learning from a country-wide assessment. PLoS ONE 16(4):e0249573. https://doi.org/10.1371/journal.pone.0249573.

Milton, S. J., J. R. Wilson, D. M. Richardson, C. L. Seymour, W. R. Dean, D. M. Iponga, and S. Proches. 2007. Invasive alien plants infiltrate bird-mediated shrub nucleation processes in arid savanna. Journal of Ecology 95:648-661. https://doi.org/10.1111/j.1365-2745.2007.01247.x.

Murcia, C., and M. R. Guariguata. 2014. La restauración ecológica en Colombia: Tendencias, necesidades y oportunidades. Documentos Ocasionales 107. CIFOR. Bogor, Indonesia. https://doi.org/10.17528/cifor/004519.

Murcia, C., M. R. Guariguata, M. Peralvo, and V. Gálmez. 2017. La restauración de bosques andinos tropicales: Avances, desafíos y perspectivas del futuro. Documentos Ocasionales 170. CIFOR. Bongor, Indonesia. https://doi.org/10.17528/cifor/006524.

Padilla, F. M., and F. I. Pugnaire. 2006. The role of nurse plants in the restoration of degraded environments. Frontiers in Ecology and the Environment 4:196-202. https://doi.org/10.1890/1540-9295(2006)004[0196:TRONPI]2.0.CO;2.

Peterson, C. J., J. J. Dosch, and W. P. Carson. 2014. Pasture succession in the Neotropics: Extending the nucleation hypothesis into a matrix discontinuity hypothesis. Oecologia 175:1325-1335. https://doi.org/10.1007/s00442-014-2986-7.

Piaia, B. B., A. P. M. Rovedder, D. Procknow, and B. Camargo. 2021. Assessment of ecological indicators in restoration by applied nucleation with different ages. Ciencia Florestal 31:1512-1534. https://doi.org/10.5902/1980509848105.

Piaia, B. B., A. P. M. Rovedder, D. Procknow, B. Camargo, M. D. Gazzola, J. P. Croda, and M. de Moraes Stefanello. 2020. Natural regeneration as an indicator of ecological restoration by applied nucleation and passive restoration. Ecological Engineering 157:105991. https://doi.org/10.1016/j.ecoleng.2020.105991.

Ramírez-Soto, A., C. R. Lucio-Palacio, R. Rodríguez-Mesa, I. Sheseña-Hernández, F. N. Farhat, B. Villa-Bonilla, L. Landa Libreros, G. Gutiérrez Sosa, O. Trujillo Santos, I. Gómez Sánchez, and E. Ruelas Inzunza. 2018a. Restoration of tropical montane cloud forests: a six-prong strategy. Restoration Ecology 26:206-211. https://doi.org/10.1111/rec.12660.

Reis, A., F. C. Bechara, M. B. de Espíndola, N. K. Vieira, and L. L. de Souza. 2003. Restoration of damaged land areas: using nucleation to improve successional processes. Natureza and Conservação 1(1):85-92.

Reis, A., F. C. Bechara, and D. R. Tres. 2010. Nucleation in tropical ecological restoration. Scientia Agricola 67:244-250. https://doi.org/10.1590/S0103-90162010000200018.

Rey-Benayas, J. M., J. M. Bullock, and A. C. Newton. 2008. Creating woodland islets to reconcile ecological restoration, conservation, and agricultural land use. Frontiers in Ecology and the Environment 6:329-336. https://doi.org/10.1890/070057.

Rojas-Botero, S., J. Solorza-Bejarano, J. Kollmann, and L. H. Teixeira. 2020. Nucleation increases understory species and functional diversity in early tropical forest restoration. Ecological Engineering 158:106031. https://doi.org/10.1016/j.ecoleng.2020.106031.

Romijn, E., R. Coppus, V. De Sy, M. Herold, R. M. Roman-Cuesta, and L. Verchot. 2019. Land Restoration in Latin America and the Caribbean: An Overview of Recent, Ongoing and Planned Restoration Initiatives and Their Potential for Climate Change Mitigation. Forests 10:510. https://doi.org/10.3390/f10060510.

Ruíz, J., O. Vargas, and N. Rodríguez. 2023. Restoration priorities: Integrating successional states and landscape resilience in tropical dry forest compensation projects in Colombia. Applied Geography 157:103021. https://doi.org/10.1016/j.apgeog.2023.103021.

Sabogal, C., C. Besacier, and D. McGuire. 2015. Restauración de bosques y paisajes: conceptos, enfoques y desafíos que plantea su ejecución. Unasylva 66(3):3-10.

Shaw, J. A., L. M. Roche, and E. S. Gornish. 2020. The use of spatially patterned methods for vegetation restoration and management across systems. Restoration Ecology 28:766–775. https://doi.org/10.1111/rec.13198.

Shimamoto, C. A., A. A. Padial, C. M. da Rosa, and M. C. M. Marques. 2018. Restoration of ecosystem services in tropical forests: A global meta-analysis. PLoS ONE 13(12):e0208523. https://doi.org/10.1371/journal.pone.0208523.

Stanturf, J. A., and S. Mansourian. 2020. Forest landscape restoration: state of play. Royal Society Open Science 7:201218. https://doi.org/10.1098/rsos.201218.

Strassburg, B. B. N., A. Iribarrem, H. L. Beyer, C. L. Cordeiro, R. Crouzeilles, et al. 2020. Global priority areas for ecosystem restoration. Nature 586(7381):724-729. https://doi.org/10.1038/s41586-020-2784-9.

Trentin, B. E., D. A. Estevan, E. F. S. Rossetto, M. R. Gorenstein, G. P. Brizola, and F. C. Bechara. 2018. Restauração florestal na mata atlântica: Passiva, nucleação e plantio de alta diversidade. Ciencia Florestal 28:160-174. https://doi.org/10.5902/1980509831647.

Tres, D. R., and A. Reis. 2007. La nucleación como propuesta para la restauración de la conectividad del paisaje. II Simposio Internacional sobre restauración ecológica 11. Cuba.

Torres-Rodríguez, S., J. E. Díaz-Triana, A. Villota, W. Gómez, and A. Avella-M. 2019. Diagnóstico ecológico, formulación e implementación de estrategias para la restauración de un bosque seco tropical interandino (Huila, Colombia). Caldasia 41(1):29-42. https://doi.org/10.15446/caldasia.v41n1.71275.

[UN] General Assembly. Transforming our world: The 2030 Agenda for Sustainable Development. In: Proceedings of the General Assembly 70th Session. 2015. New York, USA.

Vargas, O. 2011. Restauración Ecológica: Biodiversidad y Conservación. Acta biológica colombiana 16(2):221-246. URL: revistas.unal.edu.co/index.php/actabiol/article/view/19280.

Vieira, I. R., F. S. de Araújo, and R. B. Zandavalli. 2013. Shrubs promote nucleation in the Brazilian semi-arid region. Journal of Arid Environments 92:42-45. https://doi.org/10.1016/j.jaridenv.2013.01.009.

Wilson, S. J., N. S. Alexandre, K. D. Holl, J. L. Reid, R. Zahawi, D. Celentano, and L. Werden. 2021. Applied nucleation restoration guide for tropical forests. Conservation International Final Report. United States.

Yarranton, G. A., and R. G. Morrison. 1974. Spatial Dynamics of a Primary Succession: Nucleation. The Journal of Ecology 62:417-428. https://doi.org/10.2307/2258988.

Zahawi, R. A., and C. K. Augspurger. 2006. Tropical forest restoration: Tree islands as recruitment foci in degraded lands of Honduras. Ecological Applications 16:464-478. https://doi.org/10.1890/1051-0761(2006)016[0464:tfrtia]2.0.co;2.

Zahawi, R. A., K. D. Holl, R. J. Cole, and J. L. Reid. 2013. Testing applied nucleation as a strategy to facilitate tropical forest recovery. Journal of Applied Ecology 50:88-96. https://doi.org/10.1111/1365-2664.12014.

Zahawi, R. A., L. K. Werden, M. San-José, J. A. Rosales, J. Flores, and K. D. Holl. 2021. Proximity and abundance of mother trees affects recruitment patterns in a long-term tropical forest restoration study. Ecography 44:1826-1837. https://doi.org/10.1111/ecog.05907.

Zalba, S., and S. R. Ziller. 2007. Adaptive management of alien invasive species: putting the theory into practice. Natureza and Conservação 5(2):86-92.

La nucleación: Una alternativa para la restauración ecológica de bosques neotropicales

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2023-11-14

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Díaz-Triana, J. E., Vargas-Ríos, O., & Rodríguez-Eraso, N. (2023). La nucleación: Una alternativa para la restauración ecológica de bosques neotropicales. Ecología Austral, 33(3), 867–886. https://doi.org/10.25260/EA.23.33.3.0.2134